/* Zachary Zeidler — research website
   Palette and type carried over from the original design; contrast raised for readability. */

:root {
  --paper: #f8f8f4;
  --white: #ffffff;
  --wash: #eef1ea;
  --ink: #1f302a;
  --body: #333d38;
  --muted: #4f5a54;
  --green: #2e5a48;
  --green-dark: #1b3e2f;
  --green-soft: #dfe8e1;
  --line: #d6dbd2;
  --learning: #4e9e49;
  --projection: #bd9c18;
  --recall: #b54d97;
  --serif: Georgia, 'Times New Roman', serif;
  --sans: -apple-system, BlinkMacSystemFont, 'Segoe UI', Roboto, 'Helvetica Neue', sans-serif;
  --header-h: 72px;
}

* { box-sizing: border-box; }
html { scroll-padding-top: calc(var(--header-h) + 16px); scroll-behavior: smooth; }
body {
  margin: 0;
  background: var(--paper);
  color: var(--body);
  font: 18px/1.7 var(--sans);
  -webkit-font-smoothing: antialiased;
  overflow-x: hidden;
}
a { color: var(--green); text-decoration-thickness: 1px; text-underline-offset: 4px; }
a:hover { color: var(--green-dark); }
a:focus-visible, button:focus-visible, summary:focus-visible { outline: 3px solid #5f8a75; outline-offset: 4px; }
button { font: inherit; }
img { max-width: 100%; display: block; }
figure { margin: 0; }
h1, h2, h3 { color: var(--ink); font-weight: 400; margin: 0; line-height: 1.18; }
h1, h2 { font-family: var(--serif); }
h1 { font-size: clamp(40px, 4.6vw, 62px); letter-spacing: -1.5px; }
h2 { font-size: clamp(30px, 3.1vw, 42px); letter-spacing: -.6px; }
h3 { font-size: 21px; line-height: 1.35; }
p { margin: 0 0 18px; }
.wrap { width: min(1160px, calc(100% - 88px)); margin: auto; }

.eyebrow {
  font-size: 12.5px; letter-spacing: 1.6px; text-transform: uppercase;
  font-weight: 600; color: var(--green); margin: 0 0 14px;
}
.small { font-size: 15px; line-height: 1.6; color: var(--muted); }

/* Skip link */
.skip-link { position: absolute; top: -100px; left: 20px; padding: 8px 12px; background: var(--white); z-index: 50; }
.skip-link:focus { top: 10px; }

/* ---------- Sticky header ---------- */
.site-header {
  position: sticky; top: 0; z-index: 20;
  background: rgb(248 248 244 / .92);
  backdrop-filter: saturate(1.4) blur(10px);
  -webkit-backdrop-filter: saturate(1.4) blur(10px);
  border-bottom: 1px solid var(--line);
}
.header-inner { display: flex; justify-content: space-between; align-items: center; gap: 28px; min-height: var(--header-h); }
.wordmark { font-size: 19px; font-weight: 600; color: var(--ink); text-decoration: none; white-space: nowrap; line-height: 1.2; }
.wordmark span { display: block; color: var(--muted); font-size: 11.5px; letter-spacing: 1.4px; text-transform: uppercase; font-weight: 400; margin-top: 3px; }
.site-nav { display: flex; gap: 6px; align-items: center; font-size: 15px; }
.site-nav a { text-decoration: none; color: var(--body); padding: 7px 11px; border-radius: 6px; white-space: nowrap; }
.site-nav a:hover { background: var(--wash); color: var(--ink); }
.site-nav a.is-active, .site-nav a[aria-current=page] { color: var(--green); background: var(--green-soft); }
.site-nav .nav-cv { margin-left: 8px; background: var(--green); color: var(--white); font-weight: 500; padding: 7px 15px; }
.site-nav .nav-cv:hover { background: var(--green-dark); color: var(--white); }

/* ---------- Buttons & links ---------- */
/* One primary style, one secondary style, and quiet text links for everything else. */
.actions { display: flex; flex-wrap: wrap; gap: 12px; align-items: center; margin-top: 28px; }
.btn {
  display: inline-flex; align-items: center; gap: 8px;
  padding: 11px 20px; border-radius: 7px; font-size: 15.5px; font-weight: 500;
  text-decoration: none; border: 1px solid var(--green); line-height: 1.2;
}
.btn-primary { background: var(--green); color: var(--white); }
.btn-primary:hover { background: var(--green-dark); border-color: var(--green-dark); color: var(--white); }
.btn-secondary { background: transparent; color: var(--green); }
.btn-secondary:hover { background: var(--green-soft); }
.quiet-link { font-size: 15.5px; color: var(--muted); text-decoration: underline; text-decoration-color: var(--line); }
.quiet-link:hover { color: var(--green); text-decoration-color: currentColor; }
.ext::after { content: " ↗"; font-size: .9em; }

/* ---------- Hero ---------- */
.hero { display: grid; grid-template-columns: minmax(0, 1fr) 342px; gap: 80px; align-items: center; padding: 64px 0 60px; }
.market-badge {
  display: inline-flex; align-items: center; gap: 10px;
  background: var(--white); border: 1px solid var(--line); border-radius: 999px;
  padding: 7px 16px 7px 13px; font-size: 14.5px; font-weight: 500; color: var(--ink); margin-bottom: 26px;
}
.market-badge .pulse { width: 9px; height: 9px; border-radius: 50%; background: var(--learning); box-shadow: 0 0 0 4px rgb(78 158 73 / .18); }
.hero h1 { margin-bottom: 24px; }
.hero .lead { font-size: 22px; line-height: 1.5; color: var(--ink); max-width: 590px; }
.hero .sub { max-width: 600px; }
.identity { display: flex; gap: 15px; align-items: center; margin-top: 34px; }
.identity img { width: 58px; height: 58px; border-radius: 50%; object-fit: cover; object-position: 50% 43%; }
.identity p { margin: 0; font-size: 15px; line-height: 1.5; color: var(--muted); }
.identity strong { font-weight: 600; color: var(--ink); }
.histology-figure { width: 100%; max-width: 342px; justify-self: center; }
.hero-image { width: 100%; height: auto; }
figcaption { font-size: 14px; line-height: 1.55; color: var(--muted); margin-top: 12px; }
.caption-key { display: inline-flex; gap: 6px; align-items: center; margin-right: 14px; white-space: nowrap; }
.dot { width: 9px; height: 9px; border-radius: 50%; display: inline-block; }
.dot.learning { background: var(--learning); }
.dot.projection { background: var(--projection); }
.dot.recall { background: var(--recall); }

.image-button { display: block; width: 100%; padding: 0; border: 0; background: transparent; cursor: zoom-in; text-align: left; }
.image-button:hover img { outline: 1px solid var(--green); outline-offset: 3px; }

/* ---------- Highlights strip ---------- */
.highlights { display: grid; grid-template-columns: repeat(4, 1fr); border-top: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.highlight { padding: 24px 24px 24px 0; text-decoration: none; color: inherit; }
.highlight + .highlight { padding-left: 24px; border-left: 1px solid var(--line); }
.highlight .kicker { display: block; font-size: 12.5px; letter-spacing: 1.2px; text-transform: uppercase; font-weight: 600; color: var(--green); margin-bottom: 6px; }
.highlight strong { display: block; font-family: var(--serif); font-weight: 400; font-size: 20px; color: var(--ink); line-height: 1.3; margin-bottom: 4px; }
.highlight span.meta { font-size: 14.5px; color: var(--muted); line-height: 1.45; display: block; }
a.highlight:hover strong { color: var(--green); }

/* ---------- Sections ---------- */
.section { padding: 84px 0; }
.section + .section { border-top: 1px solid var(--line); }
.band { background: var(--wash); border-top: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.band + .section { border-top: 0; }
.section-head { max-width: 780px; margin-bottom: 38px; }
.section-head h2 { margin-bottom: 18px; }
.section-head p:last-child { margin-bottom: 0; }

/* ---------- Featured research (full-width moment) ---------- */
.feature { background: var(--white); border-top: 1px solid var(--line); border-bottom: 1px solid var(--line); padding: 88px 0; }
.status-tag {
  display: inline-block; font-size: 13px; font-weight: 600; letter-spacing: .3px;
  background: var(--green-soft); color: var(--green-dark); padding: 5px 11px; border-radius: 5px; margin-bottom: 18px;
}
.feature h2 { font-size: clamp(34px, 4.2vw, 56px); letter-spacing: -1.2px; margin-bottom: 24px; max-width: 900px; }
.takeaway {
  font-family: var(--serif); font-size: clamp(21px, 2vw, 26px); line-height: 1.45; color: var(--ink);
  max-width: 880px; padding-left: 22px; border-left: 3px solid var(--projection); margin-bottom: 52px;
}
.feature-grid { display: grid; grid-template-columns: minmax(0, 732px) minmax(0, 1fr); gap: 56px; align-items: start; }
.feature-figure .panel { background: var(--white); border: 1px solid var(--line); padding: 18px 10px; }
.findings { list-style: none; padding: 0; margin: 0 0 28px; counter-reset: f; }
.findings li { counter-increment: f; display: grid; grid-template-columns: 34px 1fr; gap: 10px; padding: 14px 0; border-top: 1px solid var(--line); font-size: 16.5px; line-height: 1.6; }
.findings li:last-child { border-bottom: 1px solid var(--line); }
.findings li::before { content: "0" counter(f); font-size: 13px; font-weight: 600; color: var(--green); padding-top: 3px; }
.findings strong { color: var(--ink); font-weight: 600; }
.anatomy { background: #050505; padding: 10px; display: grid; grid-template-columns: 1fr 1fr; gap: 8px; color: #f4f4f4; margin-top: 22px; }
.anatomy img { width: 100%; height: auto; }
.anatomy span { display: block; font-size: 11.5px; letter-spacing: .8px; text-transform: uppercase; padding: 5px 4px 1px; }

/* ---------- Research program pillars ---------- */
.pillars { display: grid; grid-template-columns: repeat(3, 1fr); gap: 28px; }
.pillar { background: var(--white); border: 1px solid var(--line); border-top: 3px solid var(--green); padding: 0 0 26px; display: flex; flex-direction: column; }
.pillar svg { display: block; width: 100%; height: auto; background: #fbfbf8; border-bottom: 1px solid var(--line); }
.pillar-body { padding: 22px 24px 0; display: flex; flex-direction: column; flex: 1; }
.pillar .number { font-size: 13px; font-weight: 600; letter-spacing: .4px; color: var(--green); display: block; margin-bottom: 10px; }
.pillar h3 { font-family: var(--serif); font-size: 26px; margin-bottom: 14px; letter-spacing: -.3px; }
.pillar p { font-size: 16.5px; line-height: 1.65; }
.pillar .approach { font-size: 14.5px; color: var(--muted); margin: auto 0 0; padding-top: 14px; border-top: 1px dashed var(--line); }
.relevance { margin-top: 34px; font-size: 16.5px; color: var(--muted); max-width: 820px; }

/* svg text */
.svg-label { font: 600 10.5px var(--sans); fill: #4f5a54; letter-spacing: .2px; }
.svg-label-dark { font: 600 11px var(--sans); fill: #1f302a; }

/* ---------- Earlier findings ---------- */
.prior { display: grid; grid-template-columns: 1fr 1fr; gap: 28px; }
.prior-card { border-top: 1px solid var(--line); padding-top: 22px; }
.prior-card .journal { font-size: 14px; font-weight: 600; color: var(--green); margin-bottom: 8px; }
.prior-card h3 { font-family: var(--serif); font-size: 27px; margin-bottom: 14px; }
.prior-card p { font-size: 16.5px; }

/* ---------- Expertise ---------- */
.methods { display: grid; grid-template-columns: repeat(4, 1fr); gap: 34px; }
.methods h3 { font-size: 18px; font-weight: 600; margin-bottom: 10px; }
.methods p { font-size: 16px; line-height: 1.65; margin: 0; }
.method-icon { width: 34px; height: 34px; margin-bottom: 14px; color: var(--green); }
.tags { display: flex; flex-wrap: wrap; gap: 8px; margin: 40px 0 0; padding: 0; list-style: none; }
.tags li { font-size: 14.5px; background: var(--white); border: 1px solid var(--line); border-radius: 999px; padding: 5px 13px; color: var(--body); }

/* ---------- About ---------- */
.about-grid { display: grid; grid-template-columns: 280px 1fr; gap: 64px; align-items: start; }
.portrait { width: 100%; aspect-ratio: 4/5; object-fit: cover; object-position: center 40%; }
.about-grid .bio p { max-width: 720px; }
.about-grid .bio .intro { font-size: 21px; line-height: 1.55; color: var(--ink); }
.about-cols { display: grid; grid-template-columns: 1fr 1fr; gap: 56px; margin-top: 64px; }
.about-cols h3 { font-family: var(--serif); font-size: 25px; margin-bottom: 18px; }
.timeline { list-style: none; padding: 0; margin: 0; }
.timeline li { display: grid; grid-template-columns: 104px 1fr; gap: 18px; padding: 15px 0; border-top: 1px solid var(--line); font-size: 16px; line-height: 1.5; }
.timeline li:last-child { border-bottom: 1px solid var(--line); }
.timeline .date { font-size: 14.5px; color: var(--muted); padding-top: 1px; }
.timeline strong { display: block; font-weight: 600; color: var(--ink); }
.timeline span.detail { color: var(--muted); font-size: 15px; }
.prose-block { margin-top: 56px; display: grid; grid-template-columns: 1fr 1fr; gap: 56px; }
.prose-block h3 { font-family: var(--serif); font-size: 25px; margin-bottom: 14px; }
.prose-block p { font-size: 16.5px; }

/* ---------- Contact ---------- */
.contact { display: grid; grid-template-columns: 1fr auto; gap: 40px; align-items: end; }
.contact h2 { margin-bottom: 14px; }
.contact address { font-style: normal; font-size: 16.5px; color: var(--muted); line-height: 1.6; }
.contact-links { display: flex; flex-direction: column; align-items: flex-end; gap: 14px; }
.email-link { font-family: var(--serif); font-size: 26px; text-decoration: none; border-bottom: 1px solid currentColor; }
.contact-links .row { display: flex; gap: 22px; flex-wrap: wrap; justify-content: flex-end; }

/* ---------- Footer ---------- */
.site-footer { padding: 30px 0; border-top: 1px solid var(--line); font-size: 14.5px; color: var(--muted); }
.footer-inner { display: flex; gap: 24px; justify-content: space-between; align-items: center; flex-wrap: wrap; }
.footer-links { display: flex; gap: 20px; flex-wrap: wrap; }
.footer-links a { color: var(--muted); }

/* ---------- Publications page ---------- */
.page-top { padding: 64px 0 44px; }
.page-top h1 { margin-bottom: 20px; }
.page-top .intro { font-size: 21px; line-height: 1.55; max-width: 780px; color: var(--ink); }
.page-top .actions { margin-top: 22px; }
.lead-pub { background: var(--white); border: 1px solid var(--line); border-top: 3px solid var(--projection); padding: 40px; display: grid; grid-template-columns: minmax(0, 1.15fr) minmax(0, 1fr); gap: 44px; align-items: start; }
.lead-pub h2 { font-size: clamp(28px, 2.8vw, 38px); margin-bottom: 16px; }
.lead-pub .paper-title { font-weight: 600; color: var(--ink); font-size: 17px; line-height: 1.45; margin-bottom: 6px; }
.lead-pub .summary { font-size: 17px; }
.authors { font-size: 15px; color: var(--muted); margin-bottom: 16px; line-height: 1.5; }
.authors b { color: var(--ink); font-weight: 600; }
.lead-pub .panel { border: 1px solid var(--line); padding: 14px 8px; }

.second-pub { display: grid; grid-template-columns: 180px 1fr; gap: 44px; padding: 40px 0; border-bottom: 1px solid var(--line); }
.second-pub .year { font-family: var(--serif); font-size: 32px; color: var(--ink); line-height: 1.1; }
.second-pub .venue { font-size: 15px; color: var(--muted); margin-top: 6px; }
.second-pub h2 { font-size: clamp(26px, 2.5vw, 32px); margin-bottom: 14px; }
.second-pub .paper-title { font-weight: 600; color: var(--ink); font-size: 17px; margin-bottom: 6px; }

.pub-group { padding-top: 56px; }
.pub-group > h2 { font-size: 28px; margin-bottom: 8px; }
.pub-list { list-style: none; margin: 0; padding: 0; }
.pub-item { display: grid; grid-template-columns: 90px 1fr; gap: 24px; padding: 22px 0; border-top: 1px solid var(--line); }
.pub-item:last-child { border-bottom: 1px solid var(--line); }
.pub-item .y { font-family: var(--serif); font-size: 22px; color: var(--ink); line-height: 1.3; }
.pub-item .t { font-size: 17.5px; font-weight: 600; color: var(--ink); line-height: 1.4; margin-bottom: 6px; display: block; }
.pub-item a.t { text-decoration: none; }
.pub-item a.t:hover { color: var(--green); text-decoration: underline; }
.pub-item .authors { margin-bottom: 4px; }
.pub-item .venue { font-size: 15px; color: var(--body); }
.pub-item .venue em { font-style: italic; }
.pill { display: inline-block; font-size: 12px; font-weight: 600; letter-spacing: .3px; padding: 2px 9px; border-radius: 4px; margin-left: 8px; vertical-align: 2px; background: var(--green-soft); color: var(--green-dark); }
.pill.first { background: #f3ecd2; color: #6b5508; }
.pub-item .extra { display: flex; gap: 18px; flex-wrap: wrap; margin-top: 8px; }
.note { font-size: 14.5px; color: var(--muted); margin-top: 40px; }

/* ---------- Image dialog ---------- */
.image-dialog { width: min(1120px, 94vw); max-height: 93vh; border: 0; padding: 24px; background: var(--paper); color: var(--body); }
.image-dialog::backdrop { background: rgb(16 25 20 / .82); }
.image-dialog .dialog-top { display: flex; justify-content: space-between; align-items: center; gap: 20px; margin-bottom: 18px; }
.image-dialog h2 { font-size: 24px; }
.image-dialog .close { border: 1px solid var(--line); background: var(--white); padding: 6px 14px; cursor: pointer; border-radius: 6px; }
.image-dialog img { max-height: 70vh; width: auto; max-width: 100%; height: auto; object-fit: contain; margin: auto; }
.image-dialog p { font-size: 15px; margin: 14px 0 0; }

/* ---------- Responsive ---------- */
@media (max-width: 1020px) {
  .wrap { width: calc(100% - 56px); }
  .hero { grid-template-columns: minmax(0, 1fr) minmax(0, 300px); gap: 40px; }
  .highlights { grid-template-columns: 1fr 1fr; }
  .highlight:nth-child(3) { padding-left: 0; border-left: 0; }
  .highlight:nth-child(n+3) { border-top: 1px solid var(--line); }
  .feature-grid { grid-template-columns: 1fr; gap: 36px; }
  .pillars { grid-template-columns: 1fr; max-width: 640px; }
  .methods { grid-template-columns: 1fr 1fr; gap: 30px; }
  .about-grid { grid-template-columns: 220px 1fr; gap: 40px; }
  .about-cols, .prose-block { gap: 40px; }
  .lead-pub { grid-template-columns: 1fr; padding: 32px; }
}

@media (max-width: 760px) {
  :root { --header-h: 104px; }
  body { font-size: 17px; }
  .wrap { width: calc(100% - 32px); }
  .header-inner { flex-direction: column; align-items: flex-start; gap: 8px; padding: 12px 0 8px; }
  .wordmark span { display: inline; margin-left: 8px; font-size: 10.5px; }
  .site-nav { width: 100%; overflow-x: auto; scrollbar-width: none; gap: 2px; font-size: 14.5px; margin-left: -8px; padding-bottom: 2px; }
  .site-nav::-webkit-scrollbar { display: none; }
  .site-nav a { padding: 6px 9px; }
  .site-nav .nav-cv { margin-left: 4px; padding: 6px 12px; }
  .hero { grid-template-columns: 1fr; padding: 36px 0 44px; gap: 34px; }
  .hero h1 { font-size: 42px; letter-spacing: -1px; }
  .hero .lead { font-size: 19.5px; }
  .market-badge { font-size: 13.5px; }
  .highlights { grid-template-columns: 1fr; }
  .highlight, .highlight + .highlight { padding: 18px 0; border-left: 0; }
  .highlight + .highlight { border-top: 1px solid var(--line); }
  .section { padding: 56px 0; }
  .feature { padding: 56px 0; }
  .takeaway { margin-bottom: 36px; padding-left: 16px; }
  .findings li { font-size: 16px; }
  .prior { grid-template-columns: 1fr; }
  .methods { grid-template-columns: 1fr; gap: 26px; }
  .about-grid { grid-template-columns: 1fr; gap: 28px; }
  .portrait { max-width: 220px; }
  .about-cols, .prose-block { grid-template-columns: 1fr; gap: 40px; margin-top: 44px; }
  .timeline li { grid-template-columns: 92px 1fr; gap: 12px; }
  .contact { grid-template-columns: 1fr; }
  .contact-links { align-items: flex-start; }
  .contact-links .row { justify-content: flex-start; }
  .email-link { font-size: 22px; }
  .page-top { padding: 40px 0 32px; }
  .lead-pub { padding: 22px; gap: 26px; }
  .second-pub { grid-template-columns: 1fr; gap: 12px; padding: 32px 0; }
  .second-pub .year { font-size: 26px; }
  .pub-item { grid-template-columns: 1fr; gap: 4px; }
  .pub-item .y { font-size: 18px; }
  .image-dialog { padding: 16px; }
}

@media (prefers-reduced-motion: reduce) { html { scroll-behavior: auto; } }

/* Phones: keep the nav to the four links that matter most so the CV button is always visible */
@media (max-width: 520px) {
  .site-nav a[href$="#directions"], .site-nav a[href$="#expertise"] { display: none; }
  .site-nav { overflow: visible; }
}

/* ---------- v2 layout: portrait hero, simple directions, figure row ---------- */
.hero { grid-template-columns: minmax(0, 1fr) 380px; padding: 72px 0 80px; }
.hero-portrait { width: 100%; max-width: 380px; justify-self: end; }
.hero-portrait img { width: 100%; height: auto; aspect-ratio: 4/5; object-fit: cover; object-position: center 35%; border-radius: 4px; }
.hero-portrait figcaption { margin-top: 12px; }
.pillar { padding-bottom: 26px; }
.pillar-body { padding-top: 26px; }
.figure-row { display: grid; grid-template-columns: minmax(0, 342px) minmax(0, 1fr); gap: 28px; align-items: start; margin-bottom: 44px; }
.figure-row .panel { background: var(--white); border: 1px solid var(--line); padding: 18px 10px; }
.findings-cols { display: grid; grid-template-columns: repeat(3, 1fr); gap: 0 32px; margin-bottom: 8px; }
.findings-cols li, .findings-cols li:last-child { border-bottom: 1px solid var(--line); }
.findings-cols + .actions { margin-top: 24px; }
.methods { grid-template-columns: repeat(3, 1fr); row-gap: 40px; }
.method-wide { grid-column: span 2; }
.bio.section-head { max-width: 820px; }
.bio .intro { font-size: 21px; line-height: 1.55; color: var(--ink); }
.about-cols { margin-top: 24px; }
#publications .pub-list { margin-bottom: 8px; }
@media (max-width: 1020px) {
  .hero { grid-template-columns: minmax(0, 1fr) 300px; }
  .figure-row { grid-template-columns: 1fr 2fr; }
  .methods { grid-template-columns: 1fr 1fr; }
  .method-wide { grid-column: 1 / -1; }
}
@media (max-width: 760px) {
  .hero { grid-template-columns: 1fr; padding: 32px 0 48px; }
  .hero-portrait { max-width: 240px; justify-self: start; }
  .figure-row { grid-template-columns: 1fr; }
  .figure-row figure:first-child { max-width: 342px; }
  .findings-cols { grid-template-columns: 1fr; }
  .findings-cols li { border-bottom: 0; }
  .findings-cols li:last-child { border-bottom: 1px solid var(--line); }
  .methods { grid-template-columns: 1fr; }
}
@media (max-width: 760px) {
  .wordmark span { display: block; margin: 2px 0 0; }
}

/* v3: whole-brain pair in featured row, uniform expertise cards, relevance under intro */
.figure-row { grid-template-columns: minmax(0, 342px) minmax(0, 1fr); }
.figure-row .anatomy { margin-top: 0; }
.section-head .relevance { margin-top: 16px; padding-left: 16px; border-left: 3px solid var(--green); color: var(--ink); font-size: 17px; }
.methods { grid-template-columns: repeat(3, 1fr); }
@media (max-width: 1020px) { .methods { grid-template-columns: 1fr 1fr; } }
@media (max-width: 760px) { .methods { grid-template-columns: 1fr; } }
.anatomy > div, .figure-row > figure { min-width: 0; }
.anatomy { grid-template-columns: minmax(0, 1fr) minmax(0, 1fr); }
.anatomy { overflow: hidden; }
.anatomy span { white-space: normal; overflow-wrap: anywhere; }
@media (max-width: 760px) { .anatomy span { font-size: 10px; letter-spacing: .4px; } }
@media (max-width: 1020px) { .figure-row { grid-template-columns: minmax(0, 1fr) minmax(0, 1.6fr); } }
@media (max-width: 760px) { .figure-row { grid-template-columns: minmax(0, 1fr); } }

/* v4: smaller histology panel, caption-only whole-brain panel */
.figure-row { grid-template-columns: minmax(0, 260px) minmax(0, 1fr); gap: 32px; }
.figure-row figcaption { max-width: 560px; }
@media (max-width: 1020px) { .figure-row { grid-template-columns: minmax(0, 220px) minmax(0, 1fr); } }
@media (max-width: 760px) { .figure-row { grid-template-columns: minmax(0, 1fr); } .figure-row figure:first-child { max-width: 260px; } }

/* v5: square histology sized to match the whole-brain panel height
   (panel height ≈ 26.4% of row width given two 808×579 images, 10px padding, 8px gap, 32px column gap) */
.figure-row { grid-template-columns: calc(26.4% - 1px) minmax(0, 1fr); gap: 32px; }
.figure-row figure:first-child img { width: 100%; height: auto; aspect-ratio: 1 / 1; object-fit: cover; }
@media (max-width: 1020px) { .figure-row { grid-template-columns: calc(26.4% - 1px) minmax(0, 1fr); } }
@media (max-width: 760px) { .figure-row { grid-template-columns: minmax(0, 1fr); } .figure-row figure:first-child { max-width: 240px; } }